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Oligocene Epoch (meaning "slightly recent") was the third and
youngest division of the Paleogene, and the characterized by an increasing
proportion of "modern" animals.
Plate Tectonics
There was an increase in volcanic activity, and plate tectonic movement,
as India collided with Asia. The last remnant of the supercontinent
of Gondwanaland broke up as Australia and South America both separated
from Antarctica
Climate
The Oligocene also marked the start of a generalized cooling, with glaciers
forming in Antarctica for the first time during the Cenozoic. The increase
in ice sheets led to a fall in sea level. The tropics diminished, giving
way to cooler woodlands and grasslands. Although there was a slight
warming period in the late Oligocene, the overall cooling trend was
to continue, culminating in the Ice Ages of the Pleistocene.
Plant Life
By the Oligocene the major evolution and dispersal of modern types of
angiosperms had occurred. The vegetation of the higher latitudes in
the northern hemisphere changed from an essentially broad-leaved tropical
evergreen forest such as had typified the Eocene, to a temperate deciduous
woodland of evergreen and broad-leaved trees. This type of woodland
is seen today only in certain relict areas like the North Island of
New Zealand and the tip of the South African Cape. Grasses, which appeared
for the first time as plants of water margins in the Eocene, became
more common in open habitats.
In North America the flora
consisted of a mixture of subtropical elements, such as cashews and
lychee trees, with temperate trees such as roses, beech and pine. Leguminous
plants of the pea and bean family were common, as were sedges, bulrushes
and a variety of ferns.
Animal Life
In the seas the Nummulitid
foraminifera continue with some abundance. The genus Lepidocyclina
replaces Orthophragmina. Irregular echinoids such as Scutella and
Clypeaster first appear
Following the terminal
Eocene extinction which took out the Dinocerata, Archeoceti, and
most of the Titanotheres and creodonts, new kinds of mammals evolved
and expanded in an evolutionary radiation of many new types. These
included the prehistoric ancestors of dogs, cats, rhinoceroses (including
both small slender running types and hippo-like semi-aquatic forms),
and horses (such as Mesohippus, top of page).

But the most important transition
among terrestrial mammals involved the artiodactyls (even-toed ungulates)
taking over from the perrissodactyls as the dominant medium-sized herbivores
in the middle Tertiary. These included the most common animals of the
period, the sheep-like oreodonts which flourished in huge numbers. Meanwhile
the omnivorous niches were filled by giant pig-like entelodonts like
Archaeotherium (shoulder height 1 metre), which retained short legs
and low-crowned teeth.
While artiodactyls appeared
in the late Paleocene, they remained rare, and it was only during the
Oligocene that advanced forms (the first camels) developed a rumen,
a complex fore-stomach that aids in the digestion of cellulose. This
provided these animals with a great advantage in dealing with a fibrous
diet, such as was appearing with the vegetation of the time, and would
become even more important during the rest of the Cenozoic as the world
continued to become drier and cooler.
The largest animals remained
- as in the late Eocene - the perrisodactyls. The huge hornless
rhinos (Indricotherium - weight 15 to 25 tonnes) of central Asia
was the largest land mammals ever to live, matching even the great
sauropod dinosaurs in size. These enormous beasts continued for
some fifteen million years in Asia, surviving until the Mid Miocene
Meanwhile, ancestral elephant-like
forms and the rhinoceros-like Arsinotheres grew to large size in Africa,
where the first anthropoid apes appeared as well.
In South America, completely
different herbivores evolved, the Edentates (sloths, armadillos, etc),
the strange Meridiungulates, and, in view of the absence of placental
carnivores, various lines of marsupial predators and the giant flightless
flesh-eating Phorusrhacids birds - 1.5 meters and more in height.
Little is known of life
in Australia and Antarctica at this time.
The information
of this page came from http://palaeos.com/
For more
on the Cenozoic periods go to http://palaeos.com/Cenozoic/Cenozoic.htm
Page uploaded on WebDyer Site on 10 June 2004 ,
last modified 10 June 2004
text content © M. Alan Kazlev 1998-2002
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